Inflorescences short racemose ; calyx lobes unequal , neither flabellate nor all dentate .

花序短总状；萼裂片不等长，既不扇形也不全部具牙齿。

Leaf margins sparsely serrate ; fruiting bracts flabellate , with soft , thornlike , tuberculate appendages .

叶缘疏生有锯齿；扇形的果苞片，柔软的具，刺状，具瘤附属物。

Basal leaves suborbicular or flabellate , base broadly cuneate , rounded , subtruncate , or subcordate ; flowers solitary , terminal .

基生叶近圆形的或扇形的，基部宽楔形，圆形，近截形，或近心形；花单生，顶生。